mesic to mesoxeric, from the
forest-steppe to boreal belt but also found not rarely as a ruderal
western Palearctic
Asplenietea, Androsacetalia
vandelii, Thlaspietea, Galio-Urticetea, Querco-Fagetea,
more and more found as a ruderal in Dauco-Melilotion
and Artemisietea